Who is main villain of Over the Hedge, vincent the bear or the Exterminator or Miss Glady?

Vincent the Bear, I think, makes the most sense thematically and story wise. He is RJ's former best friend and opens in the beginning of the films narrative as a bear RJ tries to steal from and will kill him/eat him if he doesn't replace his food and cart and makes a dramatic, intense and epic return in the climax. He may be the biggest threat technically, because he can physically grab them and has his claws and can just eat them in 2 seconds if he grabs them.

The Exterminator, is an obvious choice as well, since he may be the most obvious threat to the animals. His electrocuter stun thingy and the lasers he installed. But they are smart enough to evade them, making tha bear more stronger.

Miss Glady is the one we see the most of, she is also an obvious choice as well I think because she is the one they try to steal from and gets angry at the animals the most. But she is obviously not the most biggest threat to them.

Who is main villain of Over the Hedge, vincent the bear or the Exterminator or Miss Glady?
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