Girls, How do boobs hurt when running?

I know that boobs can hurt when girls run, but I never hear exactly what that means. I have a pretty muscular chest that bounces when I run, but it doesn't actually hurt-- just a little uncomfortable from time to time. Do the developed part of your breasts actually hurt when you run or is it just the pulling of the skin thats uncomfortable? I'm assuming both but I've never heard anyone actually confirm!
Girls, How do boobs hurt when running?
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